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Shorebirds
Order Charadriiformes

Shorebirds are most famous for their incredible feats of migration. Some fly non-stop for 3-4 days, equivalent to a human running continuous 4-minute miles for 60 hours.

Why do shorebirds migrate?
Actually, it is more amazing that all birds don't migrate. Having wings, it makes sense for birds to follow changes in availability of food, breeding sites and to avoid predators.

Mangrove and wetland wildlife at
Sungei Buloh Nature Park
Main features: Generally small to medium, with long slender legs, necks and beaks; wings long and narrow for fast flight.

Status in Singapore:
A large variety are visiting migrants.
World distribution: Worldwide except Antarctic.

Classification: Order Charadriiformes, a vast assembly of birds. Shorebirds make up the bulk, and includes curlews, sandpipers, snipe, oystercatchers, plovers, dotterels, thick-knees, stone curlews, ibisbill, stilts, avocets, jacanas, coursers, pratincoles, seedsnipes, sheathbills. The 3 other major groups are the gulls, terns and auks.
In the warmer south, food supplies are sufficient for subsistence but not for breeding. In the north during the summer, there is an explosion of food resources due to the long daylight hours. Combined with a lack of permanent predators due to the long winter season, the north is an ideal place for breeding. Most birds fly north to the Arctic to breed, instead of south to Antarctic, because there is more land mass in the north.

Dangers in migration: Staging areas are essential to allow birds to replenish their reserves. Before migrating, shorebirds undergo hormonal changes which induce them to lay down fat. They must double their body weight before they can embark on their migration. Their reliance on staging areas make migratory shorebirds vulnerable. Without the staging areas, they may not survive to breed.

In nature, most natural mortality of adult shorebirds occurs on the breeding grounds and is due to bad weather. Bad weather on breeding grounds also kills many eggs and young, and may sometimes wipe out breeding for the year. Even under ideal conditions, shorebirds produce few eggs (average 4) and fewer young make it the journey to the south safely. Away from the breeding grounds, mortality among adults is very low. 70-95% of adult birds usually return to breed. Shorebirds also live a long time (large birds up to 30 years, medium 10-20 years, small 4-10 years).

Females usually leave the breeding sites first. Exhausted from producing eggs, it is an advantage for them to arrive at the staging area early and have first pick of the food. Males usually leave next, and the juveniles later. These waves are not due to the lack of food on the northern summer grounds, but because there is more food at the staging area for those that arrive first.

For more about the Mysteries of Migration.

Feeding styles: Shorebirds need to be efficient feeders. They need to eat up to one-third their body weight every day to fuel their active lifestyle as well as build up fat reserves for their long migrations. In some, fat reserves can make up to 30% of their body weight.

Shorebirds can be broadly divided into either "pickers" or "probers". Pickers search for food by sight and forage in a typical run-and-peck manner. They run a long distance, up to a few metres, then abruptly stop with their heads held high, and sometimes end with a peck. Plovers use this method. They also have good night vision and they feed both at night and during the day.

Probers usually have long bills and they stick these into the soft mud or sand to feel for prey. Some "mine" the surface in a "stitching" or "sewing-machine" manner. Their bills are not rigid insensitive probes. The Sandpiper's bill tip is mobile and can act as a finger tip to grasp or grip prey. Under the horny layer, the bill is rich with tactile organs.

There are, of course, other ingenious bills. Oystercatchers have a triangular bill that is a cross between a knife and a chisel. They may use these to either stab into an open bivalve and severe the muscles that close the shells; or to smash open the shell. Shorebirds with upturned bills use it to "scythe" the water, sweeping it back and forth to stir up and snag prey.

Shorebirds have other ways to detect or find prey. Snipe are believed to be able to use their feet to sense underground prey. Many Plovers foot-tremble: lifting one foot and vibrating the mud with their toes. These apparently persuade prey to show themselves to the birds. Turnstones, as their names imply, find food by turning over seaweed and small stones.

Phalaropes have an ingenious way to gather food at their arctic breeding sites. They spin above the water at one revolution per second creating a whirlpool beneath them which whips up bottom-dwelling insect larvae. As these rise to the surface, they are picked off with the Phalaropes' needle-like beaks.

How can so many birds find food in the same place? Although huge flocks of different shorebirds may be found in one location, they divide up the territory among them. There is a horizontal division: some search above the tideline, others follow the waterline, yet others in the shallows and some in deeper water still (those with longer legs). There is also a vertical division: some pick titbits off the ground, others probe underground-some deeper than others, yet others prey on creatures living deeper in the water, and others on titbits floating on the water surface.


Breeding: Shorebirds nest on treeless habitats, mainly on the Arctic tundra. During the summer of long days, the tundra supports an explosion of insects (larvae and adults) and plant food. This allows shorebirds to quickly raise their young, which often must migrate soon after they fledge.

For example, Plover chicks hatch fully feathered and able to move about and feed on their own soon after. The parents, however, still have to protect them from predators and the elements; keeping them cool from the sun and warm when it is cold. The breeding success of Plovers depend on the population cycles of lemmings and voles. When these creatures are abundant, predators such as Arctic foxes do not turn to Plovers, their eggs and chicks.
The Mysterious Matings of Shorebirds

Most shorebirds are monogamous, although a few form lifelong pair bonds. A few are more promiscuous. Some sandpipers are polygynous: one male mating with many females, and the female incubating and raising the young on her own. Males are territorial and those with good feeding grounds attract the most females. In this way, the system makes sense.

Some shorebirds are polyandrous: one female mating with more than one male, and it is the male that incubates and raises the young, while the female defends the territory and chases off other females. (Less than 1% of birds practice this, and most are shorebirds: phalaropes, spotted sandpiper). Yet others practice sequential polyandry: the female lays one clutch that is incubated by the male, then a second clutch that she incubates herself (mountain plover and sanderling).
LINKS
  • BirdWatch Ireland: Migration-changing with the seasons: excellent explanation of migration with maps of migration in Europe and Africa; and excellent illustrations of how birds migrate plus details of experiments to highlight their talents; the effects of man on bird migration.
  • Northern Prairie Wildlife Research Center: Migration of Birds, based on the book by by Frederick C. Lincoln; Tons of details about bird migration in the Americas; origin and evolution, how scientists study bird migration, migration routes and patterns, and lots lots more.
  • The Why Files: lots of details about bird migration in the Americas; why and how they do it and threats to migratory birds.
  • Birding.About.com: lots of details about why and how birds migrate, with examples, details and maps about migration in the Americas.
  • Audubon Society: Bird Migration Facts with lots of details on why wetlands are crucial for migrating birds.
  • The National Aviary: a brief description of why birds migrate.
  • US Fish and Wildlife Service Sister Shorebird Schools Home Page: fact sheets about shorebirds in general, and some very interesting facts about shorebird behaviour.
REFERENCES
  To buy these references & others, visit
Nature's Niche

  • Harry Thurston, "The World of Shorebirds", Sierra Club Books, 1996.
 
By Ria Tan, 2001